Hello,
I have an Octavia with a VP 37 and an Eberspächer D4WSC (see signature). The auxiliary heater is mounted at the front on the left side, behind the bumper. It is integrated into the water circuit between the cylinder head (suction side of the heater) and the heat exchanger (pressure side of the heater).
Circulation direction? Let's start at the heat source and end there as well: auxiliary heater, heat exchanger, engine block, auxiliary heater.
I can't think of any plausible reason why it would be different for other vehicles based on the Golf IV platform or for Webasto products.
I hope this helps you.
Regards, mcgregg.